At the 2011 ICCA Finals held at Lincoln Center, the Barden University Bellas, a talented all-female a cappella group, face a crushing defeat when their all-male rivals, the Treblemakers, triumph after senior leader Aubrey Posen is overcome by stage fright and vomits onstage. The setback casts a shadow over the Bellas, but the story follows the ripple effects of that moment as new chapters begin to unfold.
Four months later, a new path opens for Beca Mitchell, who enrolls as a freshman at Barden because her father, a professor there, insists she try college for a year before pursuing a career as a music producer. Rather than diving straight into classwork, she spends her time crafting mash-ups of songs and interning at the campus radio station, where she quickly befriends fellow freshman Jesse Swanson. The campus buzz shifts as Aubrey and her friend Chloe Beale look to recruit fresh faces for the Bellas, though interest is tepid in the wake of the earlier embarrassment.
Beca auditions with a standout performance of Cups (When I’m Gone), earning a coveted spot in the Bellas alongside seven newcomers: the tough Cynthia Rose Adams, the bold Stacie Conrad, the quirky Lilly Onakuramara, the funny Fat Amy Hobart, Jessica Smith, Ashley Jones, and Denise. Simultaneously, Jesse joins the Treblemakers, while his roommate Benji Applebaum is turned away despite a strong audition because of his awkward demeanor. The Bellas enter the 2012 Regional competition, and at Aubrey’s insistence they perform the familiar medley that had previously carried them to the Finals. The Treblemakers take first place, with the Bellas in second and advancing to the Semi-Finals.
After regional competition, the Bellas attempt to step in and break up a fight between the Treblemakers and the Tonehangers. In the chaos, Beca and Fat Amy accidentally break a window with the Treblemakers’ trophy, leading to Beca’s arrest. Jesse mobilizes help to bail her out, but the incident strains Beca’s relationships with both Jesse and Aubrey. Back in practice, Aubrey clings to the same old setlist, insisting on sticking to the plan, while Beca champions a more daring approach.
On the way to the Semi-Finals, the Treblemakers harass Fat Amy, forcing the Bellas to travel on a shared bus after they run out of gas. For their Semi-Finals performance, Beca secretly layers an arrangement of Bulletproof into the Bellas’ opening song, The Sign, to inject energy into a bored audience. The audacious move earns praise from the crowd and judges, but Aubrey lashes out, accusing Beca of breaking the Bella code by getting involved with Jesse and deviating from the plan. When Jesse denies any romantic involvement, Beca loses her temper, snaps at both of them, and ultimately quits the Bellas.
Despite the turmoil, the Bellas still place third at the Semi-Finals. A revelation then surfaces: the leader of the second-place Footnotes is still in high school, which triggers their disqualification and clears the path for the Bellas to move forward to the Finals. During a tense rehearsal, the Bellas decide to stand up to Aubrey, and in a climactic moment, Aubrey’s distress leads to a violent vomiting episode. Beca arrives, mediates the chaos, apologizes for altering the set without approval, and asks Aubrey to allow her to rejoin. Aubrey agrees and even relinquishes half of the Bellas’ leadership to Beca, setting the stage for a more collaborative dynamic. Chloe also makes a personal breakthrough, discovering she can sing bass notes after vocal nodules surgery, adding depth to the group’s sound. Meanwhile, the Treblemakers see their own changes as Bumper Allen quits to work as a back-up singer for John Mayer, and Jesse persuades the Trebles to offer Benji a place in their lineup.
Finally, at the Finals, the Bellas deliver a show-stopping medley arranged by Beca, featuring the iconic line-up including Don’t You (Forget About Me), the theme from The Breakfast Club and a nod to one of Jesse’s favorite films. The performance becomes a powerful apology and a showcase of Beca’s creative vision, culminating in a kiss between Beca and Jesse. The Bellas triumph over the Treblemakers, clinching the national championship, and the story closes with the Bellas planning new auditions six months later, signaling a fresh era for the group.